@import url('https://fonts.googleapis.com/css2?family=DM+Sans:wght@400;500;600;700&family=Fraunces:ital,wght@0,400;0,500;0,600;1,400;1,500&display=swap');
:root{font-family:'DM Sans',sans-serif;color:#23392f;background:#f8f7ef;font-synthesis:none;font-size:16px;--green:#193e32;--gold:#edbc65;--line:#dcded2}*{box-sizing:border-box}body{margin:0}button,a,input{-webkit-tap-highlight-color:transparent}button,input{font:inherit}button{cursor:pointer}button:disabled{cursor:default;opacity:.35}button:focus-visible,a:focus-visible,input:focus-visible,summary:focus-visible{outline:3px solid #b66520;outline-offset:5px}a{color:inherit}.skip{position:absolute;left:15px;top:-90px;z-index:10;background:white;padding:15px}.skip:focus{top:10px}.masthead{max-width:1440px;margin:auto;padding:28px 5%;display:flex;align-items:center;justify-content:space-between;border-bottom:1px solid var(--line);gap:20px}.brand{text-decoration:none;font-size:25px;font-weight:700;letter-spacing:-1.1px;white-space:nowrap}.brand span{font-weight:400}.brand b{color:#b5742e;font-size:32px}.section-label{letter-spacing:2px;font-size:12px;font-weight:600}.edition{font-size:13px;color:#586a5d}main{max-width:1320px;margin:0 auto;padding:44px 5% 32px}.heading{display:flex;justify-content:space-between;align-items:flex-end;margin-bottom:32px;gap:28px}.eyebrow{font-size:12px;letter-spacing:1.9px;font-weight:700;margin:0 0 14px;color:#59735d}h1,h2,h3{font-family:'Fraunces',Georgia,serif;font-weight:400}h1{font-size:clamp(42px,5.5vw,76px);line-height:.99;letter-spacing:-2.6px;margin:0}h1 br{display:none}h1 em{color:#946329;font-weight:400}.lead{font-size:16px;line-height:1.7;color:#617064;max-width:380px;margin:0 0 3px}.explorer{display:grid;grid-template-columns:minmax(0,1.65fr) minmax(280px,1fr);border-radius:24px;overflow:hidden;border:1px solid #dfe1d5}.stage{background:#e9eddd;position:relative;min-width:0;overflow:hidden}.stage-bar{display:flex;justify-content:space-between;align-items:center;gap:10px;padding:24px 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h2{font-size:36px;line-height:1.1;letter-spacing:-.8px;margin:0 0 19px}.story-copy{font-size:16px;line-height:1.7;color:#516050;margin:0 0 22px}.fact{border-top:1px solid var(--line);padding-top:17px}.fact span{font-size:11px;letter-spacing:1.5px;font-weight:700;color:#8d6228}.fact p{font-size:14px;line-height:1.65;margin:8px 0;color:#5f6755}.step-arrows{display:flex;justify-content:space-between;align-items:center;gap:10px;margin-top:auto;padding-top:21px}.step-arrows button{border:1px solid var(--line);border-radius:50%;width:44px;height:44px;background:transparent;font-size:20px;color:var(--green)}.step-arrows span{font-size:13px}.timeline{margin:28px 0 24px}.timeline-heading{display:flex;justify-content:space-between;font-size:14px;gap:15px;margin-bottom:11px}.timeline-heading label{font-weight:600}.timeline-heading>span{color:#617064;font-size:13px}input[type=range]{width:100%;height:30px;accent-color:var(--green);margin:0;cursor:pointer}.steps{display:grid;grid-template-columns:repeat(5,1fr);gap:8px;margin-top:8px}.steps button{background:transparent;border:1px solid transparent;border-radius:13px;text-align:left;padding:12px;color:#56634e;font-size:14px;min-width:0;line-height:1.4}.steps button span{display:block;font-size:11px;margin-bottom:4px;color:#777e68}.steps small{display:block;font-size:12px;margin-top:5px;font-weight:400}.steps button[aria-pressed=true]{background:var(--green);color:#fff}.steps button[aria-pressed=true] span,.steps button[aria-pressed=true] small{color:#d4ddc9}.steps button:hover:not([aria-pressed=true]){border-color:#c4cdb6}.inside{background:#fffef8;padding:30px;border:1px solid var(--line);border-radius:20px;margin:25px 0}.inside-title h2{font-size:30px;margin:0 0 22px;line-height:1.1}.inside-title .eyebrow{margin-bottom:8px}.parts{display:flex;flex-wrap:wrap;gap:8px}.parts button{background:transparent;border:1px solid #bac7b5;padding:10px 18px;border-radius:25px;min-height:44px;font-size:14px;color:var(--green)}.parts button[aria-pressed=true]{background:var(--green);color:white;border-color:var(--green)}.part-detail{display:flex;gap:22px;padding:24px 0 0;align-items:flex-start}.part-index{font:italic 46px 'Fraunces',Georgia,serif;color:#a57635}.part-detail h3{font-size:24px;margin:0 0 7px}.part-detail p{font-size:16px;line-height:1.7;margin:0;max-width:750px;color:#56634e}.safety{display:flex;gap:16px;align-items:center;border-top:1px solid var(--line);padding-top:25px}.safety-mark{border:1px solid #b78a48;color:#8c662e;border-radius:50%;flex-shrink:0;width:32px;height:32px;text-align:center;line-height:30px;font-family:Georgia,serif}.safety p{font-size:14px;color:#616750;line-height:1.6;margin:0}.safety strong{color:#324431}.sources{margin:22px 0 0;font-size:13px;color:#617064}.sources summary{cursor:pointer;padding:10px 0;width:fit-content}.sources p{max-width:850px;line-height:1.7}.sources a{display:block;margin:12px 0;text-underline-offset:4px}footer{padding:25px 5%;display:flex;align-items:center;justify-content:space-between;gap:24px;border-top:1px solid var(--line);max-width:1440px;margin:auto}footer p,footer>span{font-size:12px;color:#65715f}footer .brand{font-size:20px}.sr-only{position:absolute;width:1px;height:1px;padding:0;margin:-1px;overflow:hidden;clip:rect(0,0,0,0);white-space:nowrap;border:0}[hidden]{display:none!important}.shell-half{transition:transform .7s ease,opacity .7s ease;transform-origin:320px 80px}.opened .shell-half{transform:translateX(75px) rotate(12deg);opacity:.18}.nest-size{transition:transform .65s ease}.inspect{transform-origin:320px 175px;transition:transform .6s ease}.zoomed .inspect{transform:translateY(5px) scale(1.3)}.inspect .focus-wall{stroke:#b36724;stroke-width:5}.comb-tier{transition:filter .2s}.focus-cells .comb-tier{filter:brightness(1.25)}
@media(min-width:1200px){.diagram{height:400px}.story{padding:36px}.story h2{font-size:40px}}@media(max-width:900px){.heading{align-items:flex-start;flex-direction:column;gap:18px}.lead br{display:none}.lead{max-width:600px}.section-label{display:none}.explorer{grid-template-columns:minmax(0,1.2fr) minmax(270px,1fr)}.story{padding:26px 22px}.story h2{font-size:32px}.diagram{height:410px}.stage-bar{padding:20px 16px}.view-label{font-size:10px}.stage-actions{flex-wrap:wrap}.steps button{padding:10px 8px}}@media(max-width:640px){.masthead{padding:18px 5%}.brand{font-size:22px}.edition{font-size:11px;max-width:115px;text-align:right}main{padding-top:28px}h1{font-size:46px;letter-spacing:-1.6px}.heading{margin-bottom:24px}.eyebrow{font-size:10px}.explorer{grid-template-columns:1fr}.diagram{height:330px}.story{padding:26px}.chapter{margin-bottom:16px}.story h2{font-size:31px}.story h2 br{display:none}.story-copy{margin-bottom:18px}.fact{padding-top:12px}.step-arrows{padding-top:16px}.steps{gap:2px}.steps button{font-size:12px;padding:10px 4px;text-align:center}.steps small{font-size:10px}.timeline-heading>span{display:none}.inside{padding:24px 19px}.inside-title h2{font-size:28px}.part-detail{gap:15px}.part-index{font-size:35px}.part-detail h3{font-size:22px}.part-detail p{font-size:15px}footer{flex-wrap:wrap;gap:8px 20px}footer>span{display:none}footer p{max-width:160px;margin-left:auto;text-align:right}}@media(prefers-reduced-motion:reduce){*,*::before,*::after{transition:none!important;animation:none!important;scroll-behavior:auto!important}}

.inside-content{display:grid;grid-template-columns:minmax(0,1fr) minmax(0,1fr);align-items:center;gap:28px;margin-top:24px}.naturalist{margin:0}.naturalist img{display:block;width:100%;height:auto;border-radius:12px}.naturalist figcaption{font-size:12px;line-height:1.5;color:#65715f;margin-top:9px}.inside-content .part-detail{padding-top:0}.diagram svg .comb-tier{pointer-events:none}@media(max-width:640px){.inside-content{grid-template-columns:1fr;gap:22px}.steps button{font-size:14px}.steps small{font-size:12px}.steps{grid-template-columns:repeat(3,1fr);gap:5px}.steps button{min-height:86px}.view-label{font-size:12px}.scale-note{font-size:12px}.eyebrow{font-size:12px}.edition{font-size:12px}.stage-bar{flex-wrap:wrap}}

/* Exploration directe et contextes saisonniers */
.stage{transition:background-color .7s ease}.stage[data-season="0"],.stage[data-season="1"]{background:#eeeadc}.stage[data-season="2"]{background:#e4ecdb}.stage[data-season="3"]{background:#eee2ca}.stage[data-season="4"]{background:#e2e8e5}.habitat{position:relative;margin:74px 20px 0;text-align:center;font-size:13px;color:#4c604c;min-height:22px}.diagram{position:relative;padding-top:0;height:300px}.diagram svg{overflow:visible}.hotspots{position:absolute;inset:0;pointer-events:none}.hotspot{position:absolute;transform:translate(-50%,-50%);width:44px;height:44px;border:2px solid #fffdf5;border-radius:50%;background:#193e32;color:#fff;font-weight:700;font-size:14px;pointer-events:auto;box-shadow:0 2px 8px #193e3230;transition:background-color .2s}.hotspot:hover,.hotspot.selected{background:#8b571f}.spot-hint{text-align:center;margin:0 18px 16px;font-size:13px;color:#4c604c}.discovery{position:relative;margin:0 20px 20px;background:#fffef8;border:1px solid #ced4c2;border-radius:14px;padding:14px 16px}.discovery-head{display:flex;justify-content:space-between;align-items:center;gap:10px}.discovery-head h3{font-size:22px;line-height:1.2;margin:0}.discovery-head button{flex-shrink:0;border:0;width:44px;height:44px;border-radius:50%;background:transparent;font-size:26px;color:#193e32}.discovery p{font-size:15px;line-height:1.6;margin:6px 0 0;color:#4d5d4b}.stage-actions{padding-top:3px}.story{min-height:475px}.stage[data-season="4"] #hi-size{filter:saturate(.45)}#hi-leaves{transition:opacity .7s}.focus-cells .comb-tier{filter:brightness(1.15)}
@media(min-width:1200px){.diagram{height:325px}}@media(max-width:900px){.diagram{height:330px}.hotspot{width:40px;height:40px}}@media(max-width:640px){.diagram{height:260px}.hotspot{width:44px;height:44px}.habitat{margin-top:72px;font-size:12px}.discovery{margin:0 14px 16px}.discovery-head h3{font-size:21px}.spot-hint{font-size:12px}.story{min-height:0}.stage-actions{gap:7px}.primary,.secondary{padding:12px 17px}.scale-note{margin-bottom:17px}.timeline{margin-top:20px}}
@media(prefers-reduced-motion:reduce){.stage,#hi-leaves{transition:none}}
.spot-leader{position:absolute;height:1px;background:#38563d;transform-origin:left center;pointer-events:none}.hotspot{z-index:1}

#detail-diagram{background:#ece9dc;border-radius:14px;overflow:hidden;padding:12px}.detail-cutaway{display:block;width:100%;height:auto;max-height:340px}.brood-legend{display:flex;flex-wrap:wrap;gap:8px 14px;font-size:12px;margin-top:12px;color:#4c604c}.brood-legend span{display:flex;align-items:center;gap:6px}.brood-legend i{display:block;width:13px;height:13px;border:1px solid #a18456;border-radius:3px}.egg-key{background:#f3dba4}.cap-key{background:#fff6d9}.empty-key{background:#715335}.focus-wall .paper-fibres{stroke-width:2;filter:contrast(1.3)}.detail-cutaway .shell-half{display:none}

/* Nid, saisons et lecture dans une même zone */
.explorer{align-items:start;background:#fffef8}.stage{padding-bottom:0}.stage .timeline{margin:14px 0 0;padding:14px 18px 17px;background:#fffef8;border-top:1px solid #d3d9ca}.stage .step-arrows{padding:0;margin:0 0 7px}.stage .step-arrows span{font-size:15px;font-weight:600}.stage .timeline-heading{margin:0;font-size:12px;color:#617064}.stage .steps{display:grid;grid-template-columns:repeat(5,minmax(0,1fr));gap:4px;margin-top:4px}.stage .steps button{font-size:13px;padding:8px 3px;min-height:44px;text-align:center;line-height:1.2;border-radius:9px}.stage input[type=range]{height:25px}.story{min-height:0;padding:27px 26px;gap:18px}.story h2{font-size:30px}.story-copy{line-height:1.6;margin-bottom:17px}.chapter{margin-bottom:16px}.story .discovery{margin:0;padding:0;border:0;border-radius:0;background:transparent}.story .discovery-head{align-items:flex-start;margin-top:15px}.story .discovery-head h3{font-size:27px;line-height:1.2;padding-top:6px}.story .discovery p{font-size:16px;line-height:1.65}.story .parts{gap:6px}.story .parts button{padding:9px 11px;font-size:13px}.compact-cutaway{margin:0;border-top:1px solid var(--line);padding-top:13px}.compact-cutaway summary{font-size:14px;cursor:pointer;padding:6px 0}.compact-cutaway .naturalist{margin-top:12px}.compact-cutaway .detail-cutaway{max-height:240px}.compact-cutaway #detail-diagram{padding:8px}.scale-note{font-size:12px;margin:11px 14px 0}.safety{margin-top:25px}.stage .diagram{height:295px}.habitat{margin-top:68px}.spot-hint{margin-bottom:12px}
@media(max-width:900px){.story{padding:22px 19px}.story h2{font-size:27px}.stage .steps button{font-size:12px}.stage .timeline{padding:12px}.story .parts button{font-size:12px;padding:8px}.stage .diagram{height:285px}}
@media(max-width:640px){.explorer{grid-template-columns:1fr}.stage .diagram{height:240px}.stage .timeline{margin-top:12px;padding:10px 13px 13px}.stage .steps{grid-template-columns:repeat(5,minmax(0,1fr))}.stage .steps button{min-height:44px;font-size:12px;padding:7px 2px}.stage .step-arrows{margin-bottom:3px}.stage .timeline-heading{font-size:12px}.story{padding:20px;gap:14px}.story h2{font-size:27px;margin-bottom:12px}.story .discovery-head h3{font-size:24px}.story .discovery p{font-size:16px;line-height:1.6}.story .parts button{font-size:14px;padding:9px 12px}.chapter{margin-bottom:10px}.habitat{margin-top:66px}.heading{margin-bottom:20px}.stage .spot-hint{font-size:13px}.compact-cutaway .detail-cutaway{max-height:220px}}

.deeper{border-top:1px solid var(--line);padding-top:12px}.deeper>button{display:flex;align-items:center;justify-content:space-between;gap:30px;min-height:44px;width:100%;border:1px solid #b7c5b0;border-radius:10px;padding:10px 14px;background:#edf1e5;color:#193e32;font:600 15px "DM Sans",sans-serif}.deeper>button:hover{background:#e1e9d8}.deeper>button span{font-size:22px;font-weight:400}#deeper-content{padding:16px 0 0}#deeper-content h3{font-size:23px;margin:0 0 10px;line-height:1.25}#deeper-content p,#deeper-content dd{font-size:15px;line-height:1.65;color:#516050;margin:0 0 12px}#deeper-content dl{margin:16px 0}#deeper-content dt{font-size:15px;font-weight:700;margin:12px 0 3px}#deeper-content dd{margin-left:0}#deeper-source{font-size:13px;text-underline-offset:3px}

.species-nav{display:flex;flex-wrap:wrap;gap:8px;max-width:1320px;margin:18px auto 0;padding:0 5%}.species-nav a{font-size:14px;text-decoration:none;border:1px solid #c3ceba;border-radius:22px;padding:9px 15px;min-height:42px;display:flex;align-items:center}.species-nav a[aria-current=page]{background:#193e32;color:#fff;border-color:#193e32}.species-choice{display:flex;align-items:center;gap:25px;margin:0 0 22px;padding:16px 20px;background:#ebeedd;border-radius:15px}.species-choice label{font-size:13px;flex:1;min-width:0}.species-choice select{display:block;margin-top:6px;width:100%;font:inherit;font-size:16px;min-height:44px;background:#fffef8;color:#23392f;border:1px solid #b8c4ad;border-radius:8px;padding:8px}.species-choice p{flex:1;font-size:14px;line-height:1.6;margin:0}.wasp-page .heading h1{font-size:clamp(40px,5vw,66px)}.wasp-page .diagram{overflow:hidden}.wasp-page .stage[data-season]{background:#eceada}.wasp-page[data-species=germanica] .paper-fibres{filter:grayscale(1)}@media(max-width:640px){.species-choice{display:block;padding:15px}.species-choice p{margin-top:12px}.species-nav{gap:6px}.species-nav a{font-size:13px;padding:8px 12px}}

/* Bourdons */
.bumble-page .opened .shell-half{transform:translateY(-20px);opacity:0;pointer-events:none}.bumble-page .diagram{overflow:hidden}.bumble-page .heading h1{font-size:clamp(40px,5vw,66px)}.bumble-page .detail-cutaway{max-height:260px}.bumble-page .zoomed .inspect{transform-origin:320px 210px;transform:scale(1.25)}

/* Galeries et rayons : deux modes de nidification */
.hornet-choice{display:flex;align-items:center;gap:18px;margin:0 0 22px;padding:15px 20px;border:1px solid #c3ceba;border-radius:14px;background:#edf0e3}.hornet-choice label{font-size:15px;font-weight:600}.hornet-choice select{font:inherit;font-size:16px;min-height:44px;flex:1;max-width:440px;min-width:0;padding:9px 12px;border:1px solid #a6b59b;border-radius:9px;background:#fffef8;color:#193e32}.hornet-choice select:focus-visible{outline:3px solid #b66520;outline-offset:3px}.polistes-page .stage[data-season="4"]{background:#e2e8e5}@media(max-width:640px){.hornet-choice{display:block;padding:14px}.hornet-choice select{display:block;width:100%;margin-top:8px}.hornet-choice label{font-size:15px}}
.bee-page .stage .steps{grid-template-columns:repeat(3,minmax(0,1fr))}
.bee-page .heading h1{font-size:clamp(38px,4.6vw,64px);overflow-wrap:normal}.bee-page .diagram{overflow:hidden}.bee-page .diagram.zoomed .drawing{transform:scale(1.2);transform-origin:center}.bee-page .hotspot{transition:none}.bee-page .stage .steps button{font-size:14px}.bee-page .view-label{font-size:12px}.bee-page .species-choice p{font-size:15px}.bee-page .detail-cutaway{max-height:280px}.carpenter-page .detail-cutaway{min-height:100px}.bee-page .spot-hint{font-size:14px}.bee-page .stage .timeline-heading{font-size:14px}.bee-page .stage .steps{gap:2px}@media(max-width:640px){.bee-page .stage .steps{grid-template-columns:repeat(3,minmax(0,1fr))}.bee-page .heading h1{font-size:40px;letter-spacing:-1.4px}.bee-page .species-choice p{font-size:15px}.bee-page .stage .diagram{height:240px}}
/* Shared reading and motion controls */
.explorer-tools{display:flex;flex-wrap:wrap;justify-content:space-between;gap:12px;margin-top:18px;padding-top:14px;border-top:1px solid #d7d6c7;font-size:13px}.explorer-tools a{color:#385341;font-weight:700}.explorer-tools label{display:flex;align-items:center;gap:8px}.explorer-tools input{width:18px;height:18px;accent-color:#385341}.reduce-motion *{animation:none!important;transition:none!important;scroll-behavior:auto!important}.mobile-season-bar{display:none}.comparison-controls{display:flex;flex-wrap:wrap;gap:18px;align-items:end;padding:20px;background:#eaece1;border-radius:16px;margin:20px 0}.comparison-controls label{display:grid;gap:8px;font-size:14px;font-weight:600}.comparison-controls select,.compare-card select{font:inherit;max-width:100%;padding:12px;border:1px solid #aeb7a6;border-radius:8px;background:#fffef9;color:#253c30;min-height:44px}.comparison-grid{display:grid;grid-template-columns:1fr 1fr;gap:22px}.compare-card{min-width:0;background:#fffef9;border:1px solid #d7d8c9;border-radius:20px;padding:24px}.compare-card svg{display:block;width:100%;height:320px}.compare-card h2{font-size:23px}.compare-card p{line-height:1.65}.compare-card select{width:100%}.compare-card a{color:#385341}.compare-intro{max-width:740px;line-height:1.6}.comparison-note{font-size:14px;line-height:1.6;color:#596253}.compare-page main{max-width:1120px;margin:auto;padding:30px 24px 64px}.compare-page h1{font-size:clamp(32px,5vw,58px)}
@media(max-width:640px){.species-nav{flex-wrap:nowrap!important;overflow-x:auto;justify-content:flex-start!important;white-space:nowrap;padding-bottom:10px}.species-nav a{flex-shrink:0}h1{font-size:34px!important;margin:12px 0!important}.lead{font-size:16px!important;line-height:1.5!important}.heading{padding-top:20px!important;padding-bottom:20px!important}.mobile-season-bar.visible{display:flex;position:fixed;bottom:0;left:0;right:0;z-index:30;gap:10px;align-items:center;padding:10px 14px calc(10px + env(safe-area-inset-bottom));background:#faf9f0;box-shadow:0 -3px 18px #263c3020}.mobile-season-bar button{width:44px;height:44px;border:1px solid #9caa96;border-radius:50%;background:#fffef9;color:#253c30;font-size:22px}.mobile-season-bar label{flex:1;font-size:11px;color:#596253}.mobile-season-bar select{display:block;width:100%;border:0;background:transparent;font-family:inherit;font-size:15px;font-weight:600;color:#253c30;min-height:28px}.mobile-season-bar button:disabled{opacity:.35}body{padding-bottom:85px}.comparison-grid{grid-template-columns:1fr}.compare-card{padding:18px}.compare-card svg{height:275px}.comparison-controls{position:sticky;top:0;z-index:3;padding:12px;gap:12px}.comparison-controls label{font-size:12px}.comparison-controls select{padding:8px}.compare-page main{padding:20px 16px 40px}}
.nest-life-controls{display:flex;align-items:center;gap:14px;max-width:560px;margin:12px auto;padding:0 20px}.nest-life-controls button{flex-shrink:0;border:1px solid #879880;border-radius:24px;padding:10px 14px;min-height:44px;background:#fafbf3;color:#254334;font-family:inherit;font-size:13px;font-weight:600;cursor:pointer}.nest-life-controls button:disabled{opacity:.6;cursor:default}.nest-life-controls p{font-size:12px;line-height:1.5;color:#53644d;margin:0}.nest-life-controls button:focus-visible{outline:3px solid #a77a30;outline-offset:3px}@media(max-width:640px){.nest-life-controls{flex-direction:column;gap:8px;text-align:center;padding:0 18px}}
.attribution-notice{max-width:1080px;margin:0 auto;padding:24px 28px 30px;border-top:1px solid #d9dece;color:#52604f;font-size:12px;line-height:1.65}.attribution-notice strong{font-size:13px;font-weight:600}.attribution-notice p{max-width:850px;margin:8px 0 0}.drawing [data-attribution],.detail-cutaway [data-attribution],.compare-drawing [data-attribution]{user-select:none;pointer-events:none}